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Nero's reign. The invasion of Parthia has been a fiasco. Two Roman legions have been brought to a stand still in the snowy mountains of Armenia, and the men are dying slowly in the cold. The rest of the Roman army is in Syria, only two weeks march away but the mountains swarm with Parthian patrols. Noreno, a half-Roman, is entrusted the mission of crossing the mountains to ask ...。艾琳（星井七濑 配音）的母亲是一名兽医，两人过着相依为命的生活。一场意外的发生令母亲和艾琳即将遭受残酷的死刑折磨，在生死存亡的紧要关头，母亲用禁断的咒术久了艾琳，自己则不幸死在了猛兽的口中。 就这样，艾琳失去了自己在这世界上的唯一一个亲人，善良的养蜂老人约翰收留了无处可去的她，将她抚养成人。一次偶然中，艾琳邂逅了在天空中自由翱翔的王兽，王兽是一种高傲的物种，绝不受人类的亲近和控制，这种高洁的性格深深的吸引了艾琳，引导着艾琳走上了和母亲一样的道路。艾琳来到了位于卡扎姆的王兽保护场，开始了学习，没想到这一选择彻底改变了她的命运。。